his work on Naval Timber and Arboriculture, published in 1831, in which he briefly but completely anticipates the theory of Natural Selection. I have ordered the book, as some passages are rather obscure, but it is certainly, I think, a complete but not developed anticipation. Erasmus always said that surely this would be shown to be the case some day. Anyhow, one may be excused in not having discovered the fact in a work on Naval Timber."[30]
De Quatrefages and De Varigny have maintained that the botanist Naudin stated the theory of evolution by natural selection in 1852. He explains very clearly the process of artificial selection, and says that in the garden we are following Nature's method. "We do not think that Nature has made her species in a different fashion from that in which we proceed ourselves in order to make our variations." But, as Darwin said, "he does not show how selection acts under nature." Similarly it must be noted in regard to several pre-Darwinian pictures of the struggle for existence (such as Herder's, who wrote in 1790 "All is in struggle ... each one for himself" and so on), that a recognition of this is only the first step in Darwinism.
Profs. E. Perrier and H. F. Osborn have called attention to a remarkable anticipation of the selection-idea which is to be found in the speculations of ��tienne Geoffroy Saint-Hilaire (1825-1828) on the evolution of modern Crocodilians from the ancient Teleosaurs. Changing environment induced changes in the respiratory system and far-reaching consequences followed. The atmosphere, acting upon the pulmonary cells, brings about "modifications which are favourable or destructive ('funestes'); these are inherited, and they influence all the rest of the organisation of the animal because if these modifications lead to injurious effects the animals which exhibit them perish and are replaced by others of a somewhat different form, a form changed so as to be adapted to (�� la convenance) the new environment."
Prof. E. B. Poulton[31] has shown that the anthropologist James Cowles Prichard (1786-1848) must be included even in spite of himself among the precursors of Darwin. In some passages of the second edition of his Researches into the Physical History of Mankind (1826), he certainly talks evolution and anticipates Prof. Weismann in denying the transmission of acquired characters. He is, however, sadly self-contradictory and his evolutionism weakens in subsequent editions--the only ones that Darwin saw. Prof. Poulton finds in Prichard's work a recognition of the operation of Natural Selection. "After inquiring how it is that 'these varieties are developed and preserved in connexion with particular climates and differences of local situation,' he gives the following very significant answer: 'One cause which tends to maintain this relation is obvious. Individuals and families, and even whole colonies perish and disappear in climates for which they are, by peculiarity of constitution, not adapted. Of this fact proofs have been already mentioned.'" Mr. Francis Darwin and Prof. A. C. Seward discuss Prichard's "anticipations" in More Letters of Charles Darwin, Vol. I. p. 43, and come to the conclusion that the evolutionary passages are entirely neutralised by others of an opposite trend. There is the same difficulty with Buffon.
Hints of the idea of Natural Selection have been detected elsewhere. James Watt,[32] for instance, has been reported as one of the anticipators (1851). But we need not prolong the inquiry further, since Darwin did not know of any anticipations until after he had published the immortal work of 1859, and since none of those who got hold of the idea made any use of it. What Darwin did was to follow the clue which Malthus gave him, to realise, first by genius and afterwards by patience, how the complex and subtle struggle for existence works out a natural selection of those organisms which vary in the direction of fitter adaptation to the conditions of their life. So much success attended his application of the Selection-formula that for a time he regarded Natural Selection as almost the sole factor in evolution, variations being pre-supposed; gradually, however, he came to recognise that there was some validity in the factors which had been emphasised by Lamarck and by Buffon, and in his well known summing up in the sixth edition of the Origin he says of the transformation of species: "This has been effected chiefly through the natural selection of numerous successive, slight, favourable variations; aided in an important manner by the inherited effects of the use and disuse of parts; and in an unimportant manner, that is, in relation to adaptive structures, whether past or present, by the direct action of external conditions, and by variations which seem to us in our ignorance to arise spontaneously."
To sum up: the idea of organic evolution, older than Aristotle, slowly developed from the stage of suggestion to
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