Biology | Page 3

Edmund Beecher Wilson
design. Such an insight we must absolutely deny to man." Still, in another place Kant admitted that the facts of comparative anatomy give us "a ray of hope, however faint, that something may be accomplished by the aid of the principle of the mechanism of nature, without which there can be no science in general." It is interesting to turn from this to the bold and aggressive assertion of Huxley: "Living matter differs from other matter in degree and not in kind, the microcosm repeats the macrocosm; and one chain of causation connects the nebulous origin of suns and planetary systems with the protoplasmic foundations of life and organization."
Do not expect me to decide where such learned doctors disagree; but I will at this point venture on one comment which may sound the key-note of this address. Perhaps we shall find that in the long run and in the large sense Kant was right; but it is certain that to-day we know very much more about the formation of the living body, whether a blade of grass or a man, than did the naturalists of Kant's time; and for better or for worse the human mind seems to be so constituted that it will continue its efforts to explain such matters, however difficult they may seem to be. But I return to our more specific inquiry with the remark that the history of physiology in the past two hundred years has been the history of a progressive restriction of the notion of a "vital force" or "vital principle" within narrower and narrower limits, until at present it may seem to many physiologists that no room for it remains within the limits of our biological philosophy. One after another the vital activities have been shown to be in greater or less degree explicable or comprehensible considered as physico-chemical operations of various degrees of complexity. Every physiologist will maintain that we cannot name one of these activities, not even thought, that is not carried on by a physical mechanism. He will maintain further that in most cases the vital actions are not merely accompanied by physico-chemical operations but actually consist of them; and he may go so far as definitely to maintain that we have no evidence that life itself can be regarded as anything more than their sum total. He is able to bring forward cogent evidence that all modes of vital activity are carried on by means of energy that is set free in protoplasm or its products by means of definite chemical processes collectively known as metabolism. When the matter is reduced to its lowest terms, life, as thus viewed, seems to have its root in chemical change; and we can understand how an eminent German physiologist offers us a definition or characterization of life that runs: "The life-process consists in the metabolism of proteids." I ask your particular attention to this definition since I now wish to contrast with it another and very different one.
I shall introduce it to your attention by asking a very simple question. We may admit that digestion, for example, is a purely chemical operation, and one that may be exactly imitated outside the living body in a glass flask. My question is, how does it come to pass that an animal has a stomach?--and, pursuing the inquiry, how does it happen that the human stomach is practically incapable of digesting cellulose, while the stomachs of some lower animals, such as the goat, readily digest this substance? The earlier naturalists, such as Linnaeus, Cuvier or Agassiz, were ready with a reply which seemed so simple, adequate and final that the plodding modern naturalist cannot repress a feeling of envy. In their view plants and animals are made as they were originally created, each according to its kind. The biologist of to-day views the matter differently; and I shall give his answer in the form in which I now and then make it to a student who may chance to ask why an insect has six legs and a spider eight, or why a yellowbird is yellow and a bluebird blue. The answer is: "For the same reason that the elephant has a trunk." I trust that a certain rugged pedagogical virtue in this reply may atone for its lack of elegance. The elephant has a trunk, as the insect has six legs, for the reason that such is the specific nature of the animal; and we may assert with a degree of probability that amounts to practical certainty that this specific nature is the outcome of a definite evolutionary process, the nature and causes of which it is our tremendous task to determine to such extent as we may be able. But this does not yet touch the most essential side of the problem.
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