the young organism at a very early stage, but in Plants they are separated at a much later date from the less isolated embryonic regions that provide for the Plant's branching; in all cases we find embryonic and germ-cells screened from the life processes of the complex organism, or taking no very obvious part in it, save to form new tissues or new organs, notably in Plants.
Again, in ourselves, and to a greater or less extent in all Animals, we find a system of special tissues set apart for the reception and storage of impressions from the outer world, and for guiding the other organs in their appropriate responses--the "Nervous System"; and when this system is ill-developed or out of gear the remaining organs work badly from lack of proper skilled guidance and co- ordination. How can we, then, speak of "memory" in a germ-cell which has been screened from the experiences of the organism, which is too simple in structure to realise them if it were exposed to them? My own answer is that we cannot form any theory on the subject, the only question is whether we have any right to INFER this "memory" from the BEHAVIOUR of living beings; and Butler, like Hering, Haeckel, and some more modern authors, has shown that the inference is a very strong presumption. Again, it is easy to over-value such complex instruments as we possess. The possessor of an up-to-date camera, well instructed in the function and manipulation of every part, but ignorant of all optics save a hand-to-mouth knowledge of the properties of his own lens, might say that a priori no picture could be taken with a cigar-box perforated by a pin-hole; and our ignorance of the mechanism of the Psychology of any organism is greater by many times than that of my supposed photographer. We know that Plants are able to do many things that can only be accounted for by ascribing to them a "psyche," and these co-ordinated enough to satisfy their needs; and yet they possess no central organ comparable to the brain, no highly specialised system for intercommunication like our nerve trunks and fibres. As Oscar Hertwig says, we are as ignorant of the mechanism of the development of the individual as we are of that of hereditary transmission of acquired characters, and the absence of such mechanism in either case is no reason for rejecting the proven fact.
However, the relations of germ and body just described led Jager, Nussbaum, Galton, Lankester, and, above all, Weismann, to the view that the germ-cells or "stirp" (Galton) were IN the body, but not OF it. Indeed, in the body and out of it, whether as reproductive cells set free, or in the developing embryo, they are regarded as forming one continuous homogeneity, in contrast to the differentiation of the body; and it is to these cells, regarded as a continuum, that the terms stirp, germ-plasm, are especially applied. Yet on this view, so eagerly advocated by its supporters, we have to substitute for the hypothesis of memory, which they declare to have no real meaning here, the far more fantastic hypotheses of Weismann: by these they explain the process of differentiation in the young embryo into new germ and body; and in the young body the differentiation of its cells, each in due time and place, into the varied tissue cells and organs. Such views might perhaps be acceptable if it could be shown that over each cell-division there presided a wise all-guiding genie of transcending intellect, to which Clerk-Maxwell's sorting demons were mere infants. Yet these views have so enchanted many distinguished biologists, that in dealing with the subject they have actually ignored the existence of equally able workers who hesitate to share the extremest of their views. The phenomenon is one well known in hypnotic practice. So long as the non-Weismannians deal with matters outside this discussion, their existence and their work is rated at its just value; but any work of theirs on this point so affects the orthodox Weismannite (whether he accept this label or reject it does not matter), that for the time being their existence and the good work they have done are alike non-existent. {0e}
Butler founded no school, and wished to found none. He desired that what was true in his work should prevail, and he looked forward calmly to the time when the recognition of that truth and of his share in advancing it should give him in the lives of others that immortality for which alone he craved.
Lamarckian views have never lacked defenders here and in America. Of the English, Herbert Spencer, who however, was averse to the vitalistic attitude, Vines and Henslow among botanists, Cunningham among zoologists, have always resisted Weismannism; but, I think, none of these
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