the second, is the "Theory of Descent."
But this theory of descent leads us at once into another dilemma. If the species originated by descent from the most closely related lower species, and under certain circumstances also from species of the same rank, and even by degeneration from the next higher, it must have occurred in one of two ways: either by leaps--called by naturalists "metamorphosis of germs" or "heterogenetic conception"--or by a succession of imperceptibly small alterations of the individuals from generation to generation. Each of these changes would have been no greater than the differences we observe to-day between the individuals of the very same species, but became in the course of time so massed and strengthened in one direction that new species have been evolved. This hypothesis is called the "Theory of Development," {26} or "Evolution." We retain this name, although well aware of the fact that the authors do not agree in their use of the term "evolution." Professor Wigand, who adopts only the theory of a descent from one primordial cell to another, and who positively rejects the idea of a progress from one fully developed species to another, claims among other things that one value of his own theory is that he secures for the idea of evolution its full meaning. The expression still has a meaning for those who reject the real descent of the species or their primordial germs one from another, and acknowledge only the ideal bond of a common plan in their successive manifestations. But as soon as we examine more closely the literal and logical meaning of the word, we shall find it of most weight when we understand by it the before-mentioned gradual evolution in opposition to the theory of progress by leaps or new creations. Moreover, it is well known that long before this no other term than evolution was used to designate the growth of a single organic individual from the primordial cell and egg to its fully developed form and vital function. Besides, we find ourselves also in harmony with most of the authors, so far as they have distinct conceptions of the different scientific problems, if we use the term "theory of evolution" for the gradual development of one species from another, in opposition to the hypothesis of a metamorphosis of germs, or even of a genealogy of primordial cells.
But each evolution theory leads again to new theories, as soon as it has to be proved in a scientific way. For it can claim a scientific worth only when sustained by earnest attempts to find and prove the {27} productive power, agencies and laws of such an evolution of species. Those attempts can be made in various ways. As a philosophical question, many attempts at solution have been made, both in ancient and modern times; but being mainly in the realm of metaphysics, they do not come within the limits of our scientific essay. As a question for investigators of natural phenomena, only two attempts of sufficient importance to be mentioned have been made. The first one was made by Lamarck, who, taking the really different ideas of descent and evolution as one, made use of the hypothesis of transmutation; thus becoming the pioneer of Darwinism. The other attempt was made by Darwin in his theory of natural selection, or struggle, for existence, and is called the "Theory of Selection."
In defining our problem, therefore, we find ourselves under the influence of a scientific law of development. The simple problem which we started from has developed into a trinity of problems and attempts at solution. The simple question of the origin of species led us into the dilemma of a generatio ?quivoca, or a descent; the hypothesis of a descent led to the dilemma of a heterogenetic conception, or an evolution; and the hypothesis of an evolution rendered necessary the attempt at explaining this evolution, and showed Darwin's method of explaining it by his selection theory. It will be well for the reader to keep distinctly in mind the difference between these problems and theories, in following our investigations, even if we cannot arrange our historical sketch according to the natural principle of division arising from these differences.
For it lies in the nature of the question itself, that {28} these theories, in their historical progress, did not appear singly, but together. Those who inclined to the theory of a descent of species could claim for it the attention of scientific investigators only after having also made the attempt at conceiving this descent in a concrete way, and according to certain analogies of observation. The only analogy of the kind appeared in the sphere of individual development and individual differences on the one hand, and in that of closely related characters of allied species on the other;
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