been probably developed? Truly, I think that the opponents of modification and variation had better leave the argument of recurrence alone, or it may prove altogether too strong for them.
To sum up,--the evidence as far as we have gone is against the argument as to any limit to divergences, so far as structure is concerned; and in favour of a physiological limitation. By selective breeding we can produce structural divergences as great as those of species, but we cannot produce equal physiological divergences. For the present I leave the question there.
Now, the next problem that lies before us--and it is an extremely important one--is this: Does this selective breeding occur in nature? Because, if there is no proof of it, all that I have been telling you goes for nothing in accounting for the origin of species. Are natural causes competent to play the part of selection in perpetuating varieties? Here we labour under very great difficulties. In the last lecture I had occasion to point out to you the extreme difficulty of obtaining evidence even of the first origin of those varieties which we know to have occurred in domesticated animals. I told you, that almost always the origin of these varieties is overlooked, so that I could only produce two of three cases, as that of Gratio Kelleia and of the Ancon sheep. People forget, or do not take notice of them until they come to have a prominence; and if that is true of artificial cases, under our own eyes, and in animals in our own care, how much more difficult it must be to have at first hand good evidence of the origin of varieties in nature! Indeed, I do not know that it is possible by direct evidence to prove the origin of a variety in nature, or to prove selective breeding; but I will tell you what we can prove--and this comes to the same thing--that varieties exist in nature within the limits of species, and, what is more, that when a variety has come into existence in nature, there are natural causes and conditions, which are amply competent to play the part of a selective breeder; and although that is not quite the evidence that one would like to have--though it is not direct testimony--yet it is exceeding good and exceedingly powerful evidence in its way.
As to the first point, of varieties existing among natural species, I might appeal to the universal experience of every naturalist, and of any person who has ever turned any attention at all to the characteristics of plants and animals in a state of nature; but I may as well take a few definite cases, and I will begin with Man himself.
I am one of those who believe that, at present, there is no evidence whatever for saying, that mankind sprang originally from any more than a single pair; I must say, that I cannot see any good ground whatever, or even any tenable sort of evidence, for believing that there is more than one species of Man. Nevertheless, as you know, just as there are numbers of varieties in animals, so there are remarkable varieties of men. I speak not merely of those broad and distinct variations which you see at a glance. Everybody, of course, knows the difference between a Negro and a white man, and can tell a Chinaman from an Englishman. They each have peculiar characteristics of colour and physiognomy; but you must recollect that the characters of these races go very far deeper--they extend to the bony structure, and to the characters of that most important of all organs to us--the brain; so that, among men belonging to different races, or even within the same race, one man shall have a brain a third, or half, or even seventy per cent. bigger than another; and if you take the whole range of human brains, you will find a variation in some cases of a hundred per cent. Apart from these variations in the size of the brain, the characters of the skull vary. Thus if I draw the figures of a Mongul and of a Negro head on the blackboard, in the case of the last the breadth would be about seven-tenths, and in the other it would be nine-tenths of the total length. So that you see there is abundant evidence of variation among men in their natural condition. And if you turn to other animals there is just the same thing. The fox, for example, which has a very large geographical distribution all over Europe, and parts of Asia, and on the American Continent, varies greatly. There are mostly large foxes in the North, and smaller ones in the South. In Germany alone, the foresters reckon some eight different sorts.
Of the
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