that the few seeds or germs of American species cast up upon the shore from time to time were mostly far too old and water-logged to show signs of life in such ungenial conditions. It was from the nearer coasts of Europe, on the contrary, that our earliest colonists seemed to come. Though the prevalent winds set from the west, more violent storms reached us occasionally from the eastward direction; and these, blowing from Europe, which lay so much closer to our group, were far more likely to bring with them by waves or wind some waifs and strays of the European fauna and flora.
I well remember the first of these great storms that produced any distinct impression on my islands. The plants that followed in its wake were a few small ferns, whose light spores were more readily carried on the breeze than any regular seeds of flowering plants. For a month or two nothing very marked occurred in the way of change, but slowly the spores rooted, and soon produced a small crop of ferns, which, finding the ground unoccupied, spread when once fairly started with extraordinary rapidity, till they covered all the suitable positions throughout the islands.
For the most part, however, additions to the flora, and still more to the fauna, were very gradually made; so much so that most of the species now found in the group did not arrive there till after the end of the Glacial epoch, and belong essentially to the modern European assemblage of plants and animals. This was partly because the islands themselves were surrounded by pack-ice during that chilly period, which interrupted for a time the course of my experiment. It was interesting, too, after the ice cleared away, to note what kinds could manage by stray accidents to cross the ocean with a fair chance of sprouting or hatching out on the new soil, and which were totally unable by original constitution to survive the ordeal of immersion in the sea. For instance, I looked anxiously at first for the arrival of some casual acorn or some floating filbert, which might stock my islands with waving greenery of oaks and hazel bushes. But I gradually discovered, in the course of a few centuries, that these heavy nuts never floated securely so far as the outskirts of my little archipelago; and that consequently no chestnuts, apple trees, beeches, alders, larches, or pines ever came to diversify my island valleys. The seeds that did really reach us from time to time belonged rather to one or other of four special classes. Either they were very small and light, like the spores of ferns, fungi, and club-mosses; or they were winged and feathery, like dandelion and thistle-down; or they were the stones of fruits that are eaten by birds, like rose-hips and hawthorn; or they were chaffy grains, enclosed in papery scales, like grasses and sedges, of a kind well adapted to be readily borne on the surface of the water. In all these ways new plants did really get wafted by slow degrees to the islands; and if they were of kinds adapted to the climate they grew and flourished, living down the first growth of ferns and flowerless herbs in the rich valleys.
The time which it took to people my archipelago with these various plants was, of course, when judged by your human standards, immensely long, as often the group received only a single new addition in the lapse of two or three centuries. But I noticed one very curious result of this haphazard and lengthy mode of stocking the country: some of the plants which arrived the earliest, having the coast all clear to themselves, free from the fierce competition to which they had always been exposed on the mainland of Europe, began to sport a great deal in various directions, and being acted upon here by new conditions, soon assumed under stress of natural selection totally distinct specific forms. (You see, I have quite mastered your best modern scientific vocabulary.) For instance, there were at first no insects of any sort on the islands; and so those plants which in Europe depended for their fertilisation upon bees or butterflies had here either to adapt themselves somehow to the wind as a carrier of their pollen or else to die out for want of crossing. Again, the number of enemies being reduced to a minimum, these early plants tended to lose various defences or protections they had acquired on the mainland against slugs or ants, and so to become different in a corresponding degree from their European ancestors. The consequence was that by the time you men first discovered the archipelago no fewer than forty kinds of plants had so far diverged from the parent forms in Europe or
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