at once among the caves of our archipelago, where it hawks to this day after our flies and beetles. This seemed to me to show very conspicuously the advantage which winged animals have in the matter of cosmopolitan dispersion; for while it was quite impossible for rats, mice, or squirrels to cross the intervening belt of three hundred leagues of sea, their little winged relation, the flitter-mouse, made the journey across quite safely on his own leathery vans, and with no greater difficulty than a swallow or a wood-pigeon.
The insects of my archipelago tell very much the same story as the birds and the plants. Here, too, winged species have stood at a great advantage. To be sure, the earliest butterflies and bees that arrived in the fern-clad period were starved for want of honey; but as soon as the valleys began to be thickly tangled with composites, harebells, and sweet-scented myrtle bushes, these nectar-eating insects established themselves successfully, and kept their breed true by occasional crosses with fresh arrivals blown to sea afterwards. The development of the beetles I watched with far greater interest, as they assumed fresh forms much more rapidly under their new conditions of restricted food and limited enemies. Many kinds I observed which came originally from Europe, sometimes in the larval state, sometimes in the egg, and sometimes flying as full-grown insects before the blast of the angry tempest. Several of these changed their features rapidly after their arrival in the islands, producing at first divergent varieties, and finally, by dint of selection, acting in various ways, through climate, food, or enemies, on these nascent forms, evolving into stable and well-adapted species. But I noticed three cases where bits of driftwood thrown up from South America on the western coasts contained the eggs or larvae of American beetles, while several others were driven ashore from the Canaries or Madeira; and in one instance even a small insect, belonging to a type now confined to Madagascar, found its way safely by sea to this remote spot, where, being a female with eggs, it succeeded in establishing a flourishing colony. I believe, however, that at the time of its arrival it still existed on the African continent, but becoming extinct there under stress of competition with higher forms, it now survives only in these two widely separated insular areas.
It was an endless amusement to me during those long centuries, while I devoted myself entirely to the task of watching my fauna and flora develop itself, to look out from day to day for any chance arrival by wind or waves, and to follow the course of its subsequent vicissitudes and evolution. In a great many cases, especially at first, the new-comer found no niche ready for it in the established order of things on the islands, and was fain at last, after a hard struggle, to retire for ever from the unequal contest. But often enough, too, he made a gallant fight for it, and, adapting himself rapidly to his new environment, changed his form and habits with surprising facility. For natural selection, I found, is a hard schoolmaster. If you happen to fit your place in the world, you live and thrive, but if you don't happen to fit it, to the wall with you without quarter. Thus sometimes I would see a small canary beetle quickly take to new food and new modes of life on my islands under my very eyes, so that in a century or so I judged him myself worthy of the distinction of a separate species; while in another case, I remember, a south European weevil evolved before long into something so wholly different from his former self that a systematic entomologist would have been forced to enrol him in a distinct genus. I often wish now that I had kept a regular collection of all the intermediate forms, to present as an illustrative series to one of your human museums; but in those days, of course, we none of us imagined anybody but ourselves would ever take an interest in these problems of the development of life, and we let the chance slide till it was too late to recover it.
Naturally, during all these ages changes of other sorts were going on in my islands--elevations and subsidences, separations and reunions, which helped to modify the life of the group considerably. Indeed, volcanic action was constantly at work altering the shapes and sizes of the different rocky mountain-tops, and bringing now one, now another, into closer relations than before with its neighbours. Why, as recently as 1811 (a date which is so fresh in my memory that I could hardly forget it) a new island was suddenly formed by submarine eruption off the coast of St. Michael's, to which the
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