Evolution in Modern Thought | Page 5

that some of these pioneers of evolutionism did more than apply the evolution-idea as a modal formula of becoming, they began to inquire into the factors in the process. Thus there were pre-Darwinian theories of evolution, and to these we must now briefly refer.[15]
In all biological thinking we have to work with the categories Organism--Function--Environment, and theories of evolution may be classified in relation to these. To some it has always seemed that the fundamental fact is the living organism,--a creative agent, a striving will, a changeful Proteus, selecting its environment, adjusting itself to it, self-differentiating and self-adaptive. The necessity of recognising the importance of the organism is admitted by all Darwinians who start with inborn variations, but it is open to question whether the whole truth of what we might call the Goethian position is exhausted in the postulate of inherent variability.
To others it has always seemed that the emphasis should be laid on Function,--on use and disuse, on doing and not doing. Practice makes perfect; c'est �� force de forger qu'on devient forgeron. This is one of the fundamental ideas of Lamarckism; to some extent it met with Darwin's approval; and it finds many supporters to-day. One of the ablest of these--Mr. Francis Darwin--has recently given strong reasons for combining a modernised Lamarckism with what we usually regard as sound Darwinism.[16]
To others it has always seemed that the emphasis should be laid on the Environment, which wakes the organism to action, prompts it to change, makes dints upon it, moulds it, prunes it, and finally, perhaps, kills it. It is again impossible to doubt that there is truth in this view, for even if environmentally induced "modifications" be not transmissible, environmentally induced "variations" are; and even if the direct influence of the environment be less important than many enthusiastic supporters of this view--may we call them Buffonians--think, there remains the indirect influence which Darwinians in part rely on,--the eliminative process. Even if the extreme view be held that the only form of discriminate elimination that counts is inter-organismal competition, this might be included under the rubric of the animate environment.
In many passages Buffon[17] definitely suggested that environmental influences--especially of climate and food--were directly productive of changes in organisms, but he did not discuss the question of the transmissibility of the modifications so induced, and it is difficult to gather from his inconsistent writings what extent of transformation he really believed in. Prof. Osborn says of Buffon: "The struggle for existence, the elimination of the least-perfected species, the contest between the fecundity of certain species and their constant destruction, are all clearly expressed in various passages." He quotes two of these:[18]
"Le cours ordinaire de la nature vivante, est en g��n��ral toujours constant, toujours le m��me; son mouvement, toujours r��gulier, roule sur deux points in��branlables: l'un, la f��condit�� sans bornes donn��e �� toutes les esp��ces; l'autre, les obstacles sans nombre qui r��duisent cette f��condit�� �� une mesure d��termin��e et ne laissent en tout temps qu'�� peu pr��s la m��me quantit�� d'individus de chaque esp��ce" ... "Les esp��ces les moins parfaites, les plus d��licates, les plus pesantes, les moins agissantes, les moins arm��es, etc., ont d��j�� disparu ou dispara?tront.".
Erasmus Darwin[19] had a firm grip of the "idea of the gradual formation and improvement of the Animal world," and he had his theory of the process. No sentence is more characteristic than this: "All animals undergo transformations which are in part produced by their own exertions, in response to pleasures and pains, and many of these acquired forms or propensities are transmitted to their posterity." This is Lamarckism before Lamarck, as his grandson pointed out. His central idea is that wants stimulate efforts and that these result in improvements which subsequent generations make better still. He realised something of the struggle for existence and even pointed out that this advantageously checks the rapid multiplication. "As Dr. Krause points out, Darwin just misses the connection between this struggle and the Survival of the Fittest."[20]
Lamarck[21] (1744-1829) seems to have thought out his theory of evolution without any knowledge of Erasmus Darwin's which it closely resembled. The central idea of his theory was the cumulative inheritance of functional modifications. "Changes in environment bring about changes in the habits of animals. Changes in their wants necessarily bring about parallel changes in their habits. If new wants become constant or very lasting, they form new habits, the new habits involve the use of new parts, or a different use of old parts, which results finally in the production of new organs and the modification of old ones." He differed from Buffon in not attaching importance, as far as animals are concerned, to the direct influence of the environment, "for environment can effect no direct change whatever upon the organisation of animals," but in regard to plants he