Studying the facts and phenomena in reference to proximate causes, and endeavoring to trace back the series of cause and effect as far as possible, Darwin's aim and processes are strictly scientific, and his endeavor, whether successful or futile, must be regarded as a legitimate attempt to extend the domain of natural or physical science. For, though it well may be that "organic forms have no physical or secondary cause," yet this can be proved only indirectly, by the failure of every attempt to refer the phenomena in question to causal laws. But, however originated, and whatever be thought of Mr. Darwin's arduous undertaking in this respect, it is certain that plants and animals are subject from their birth to physical influences, to which they have to accommodate themselves as they can. How literally they are "born to trouble," and how incessant and severe the struggle for life generally is, the present volume graphically describes. Few will deny that such influences must have gravely affected the range and the association of individuals and species on the earth's surface. Mr. Darwin thinks that, acting upon an inherent predisposition to vary, they have sufficed even to modify the species themselves and produce the present diversity. Mr. Agassiz believes that they have not even affected the geographical range and the actual association of species, still less their forms; but that every adaptation of species to climate, and of species to species, is as aboriginal, and therefore as inexplicable, as are the organic forms themselves.
Who shall decide between such extreme views so ably maintained on either hand, and say how much of truth there may be in each? The present reviewer has not the presumption to undertake such a task. Having no prepossession in favor of naturalistic theories, but struck with the eminent ability of Mr. Darwin's work, and charmed with its fairness, our humbler duty will be performed if, laying aside prejudice as much as we can, we shall succeed in giving a fair account of its method and argument, offering by the way a few suggestions, such as might occur to any naturalist of an inquiring mind. An editorial character for this article must in justice be disclaimed. The plural pronoun is employed not to give editorial weight, but to avoid even the appearance of egotism, and also the circumlocution which attends a rigorous adherence to the impersonal style.
We have contrasted these two extremely divergent theories, in their broad statements. It must not be inferred that they have no points nor ultimate results in common.
In the first place, they practically agree in upsetting, each in its own way, the generally-received definition of species, and in sweeping away the ground of their objective existence in Nature. The orthodox conception of species is that of lineal descent: all the descendants of a common parent, and no other, constitute a species; they have a certain identity because of their descent, by which they are supposed to be recognizable. So naturalists had a distinct idea of what they meant by the term species, and a practical rule, which was hardly the less useful because difficult to apply in many cases, and because its application was indirect: that is, the community of origin had to be inferred from the likeness; such degree of similarity, and such only, being held to be con-specific as could be shown or reasonably inferred to be compatible with a common origin. And the usual concurrence of the whole body of naturalists (having the same data before them) as to what forms are species attests the value of the rule, and also indicates some real foundation for it in Nature. But if species were created in numberless individuals over broad spaces of territory, these individuals are connected only in idea, and species differ from varieties on the one hand, and from genera, tribes, etc., on the other, only in degree; and no obvious natural reason remains for fixing upon this or that degree as specific, at least no natural standard, by which the opinions of different naturalists may be correlated. Species upon this view are enduring, but subjective and ideal. Any three or more of the human races, for example, are species or not species, according to the bent of the naturalist's mind. Darwin's theory brings us the other way to the same result. In his view, not only all the individuals of a species are descendants of a common parent, but of all the related species also. Affinity, relationship, all the terms which naturalists use figuratively to express an underived, unexplained resemblance among species, have a literal meaning upon Darwin's system, which they little suspected, namely, that of inheritance. Varieties are the latest offshoots of the genealogical tree in "an unlineal" order; species, those of an earlier date, but of no definite
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