it is above the seed vessel, which is, of course, a portion of the flower itself. It is a hollow tube, and contains the long style or connection between the seed vessel and the stigma, a (Fig. 2). This tube, then, must be the calyx, and the small scattered scale-like bodies, b (Fig. 2), which clothe the outside, are really calyx lobes.
[Illustration: FIG. 2.--FLOWER OF PHYLLOCACTUS, CUT LENGTHWISE.
a, Calyx Tube. b, Calyx Lobes. c, Ditto, assuming the form of Petals. d, Stamens. e, Style. f, Ovary or Seed Vessel.]
Nearer the top of the flower, these calyx lobes are better developed, until, surrounding the corolla, we find them assuming the form and appearance of petals, c (Fig. 2). The corolla is composed of a large number of long strap-shaped pointed petals, very thin and delicate, often beautifully coloured, and generally spreading outwards. Springing from the bases of these petals, we find the stamens, d (Fig. 2), a great number of them, forming a bunch of threads unequal in length, and bearing on their tips the hay-seed-like anthers, which are attached to the threads by one of their points. The style is a long cylindrical body, e (Fig. 2), which stretches from the ovary to the top of the flower, where it splits into a head of spreading linear rays, ? in. in length. When the flower withers, the seed vessel, f (Fig. 2), remains on the plant and expands into a large succulent fruit, inside which is a mass of pulpy matter, inclosing the numerous, small, black, bony seeds.
It must not be supposed that all the genera into which Cactuses are divided are characterised by large flowers such as would render their study as easy as the genus taken as an illustration. In some, such for instance as the Rhipsalis, the flowers are small, and therefore less easy to dissect than those of Phyllocactus.
The stems of Cactuses show a very wide range of variation in size, in form, and in structure. In size, we have the colossal Cereus giganteus, whose straight stems when old are as firm as iron, and rise with many ascending arms or rear their tall leafless trunks like ships' masts to a height of 60 ft. or 70 ft. From this we descend through a multitude of various shapes and sizes to the tiny tufted Mamillarias, no larger than a lady's thimble, or the creeping Rhipsalis, which lies along the hard ground on which it grows, and looks like hairy caterpillars. In form, the variety is very remarkable. We have the Mistletoe Cactus, with the appearance of a bunch of Mistletoe, berries and all; the Thimble Cactus; the Dumpling Cactus; the Melon Cactus; the Turk's cap Cactus; the Rat's-tail Cactus; the Hedgehog Cactus; all having a resemblance to the things whose names they bear. Then there is the Indian Fig, with branches like battledores, joined by their ends; the Epiphyllum and Phyllocactus, with flattened leaf-like stems; the columnar spiny Cereus, with deeply channelled stems and the appearance of immense candelabra. Totally devoid of leaves, and often skeleton-like in appearance, these plants have a strange look about them, which is suggestive of some fossilised forms of vegetation belonging to the past ages of the mastodon, the elk, and the dodo, rather than to the living things of to-day.
By far the greater part of the species of Cactuses belong to the group with tall or elongated stems. "It is worthy of remark that as the stems advance in age the angles fill up, or the articulations disappear, in consequence of the slow growth of the woody axis and the gradual development of the cellular substance; so that, at the end of a number of years, all the branches of Cactuses, however angular or compressed they originally may have been, become trunks that are either perfectly cylindrical, or which have scarcely any visible angles."
A second large group is that of which the Melon and Hedgehog Cactuses are good representatives, which have sphere-shaped stems, covered with stout spines. We have hitherto spoken of the Cactuses as being without leaves, but this is only true of them when in an old or fully-developed state. On many of the stems we find upon their surface, or angles, small tubercles, which, when young, bear tiny scale-like leaves. These, however, soon wither and fall off, so that, to all appearance, leaves are never present on these plants. There is one exception, however, in the Barbadoes Gooseberry (Pereskia), which bears true and persistent leaves; but these may be considered anomalous in the order.
The term "succulent" is applied to Cactuses because of the large proportion of cellular tissue, i.e., flesh, of their stems, as compared with the woody portion. In some of them, when young, the woody system appears to be altogether absent, and they have the
Continue reading on your phone by scaning this QR Code
Tip: The current page has been bookmarked automatically. If you wish to continue reading later, just open the
Dertz Homepage, and click on the 'continue reading' link at the bottom of the page.